This page shows some more photos of the specimen on the first page about this beetle. The portrait above clearly shows the lip and jaw antennae and the long antennae. These probably indicate the beetle has a good sense of smell underwater: Balfour-Browne (1922) observed how squeaker beetles reacted immediately when food was carefully placed in the tank water, without them seeing it. He also observed that when the beetles scrambled around on the muddy bottom, they sometimes stopped abruptly, then dived into the mud, usually catching prey. Below is the right antenna.

The first part near the head, the scapus , has many small pits. These are also found in a large part of the beetle's shell; see the photo at the bottom of the page.

Like some species of aquatic beetles, the males have enlarged tarsi on their front and middle legs. These serve to provide better grip during mating. They are covered in adhesive hairs (Dettner 1997). Below is the right front leg. Note the large spurs. These are found on all legs, as are swimming hairs.

Below is the right hind leg, also with large spines. These likely help the beetle walk on dry land and dig in the soil.

Finally, a photo showing the pits. To the right of them is the (right) large, convex compound eye.